Figure 2

Harpalinae phylogeny dated using a Bayesian relaxed clock method with all-data-combined in BEAST. Branches are proportional to time in millions of years. 95% confidence intervals for the ages of basal branches in the tree and major harpaline lineages (tribes) are indicated with blue bars. The capital letters indicate the major lineages of harpalines (see Table 3), and the Greek symbols represent the fossil calibration points used in the molecular dating analyses (see Table 1).