Fossil | Min. age for lineage (Mybp) | Period | Location | Reference | |
---|---|---|---|---|---|
α | Carabidae | 155 - 160 | Late Jurassic | South Kazakhstan and Bavaria | |
β | Abacetus | 44.1 | Middle Eocene | Baltic | |
γ | Harpalinae | 89.0 - 93.5 | Late Cretaceous | South Kazakhstan | [31] |
δ | Agonum | 49.0 - 52.0 | Early Eocene | Green River CO | [89] |
ε | Amara | 44.1 | Middle Eocene | Baltic | [86] |
ζ | Cymindoidea | 44.1 | Middle Eocene | Baltic | [86] |
η | Apristus | 44.1 | Middle Eocene | Baltic | [86] |
θ | Brachinus | 34.9 - 38.0 | Late Eocene | Florissant CO | |
ι | Bradycellus | 44.1 | Middle Eocene | Baltic | [86] |
κ | Calathus | 44.1 | Middle Eocene | Baltic | [86] |
λ | Diplocheila | 34.9 - 38.0 | Late Eocene | Florissant CO | |
μ | Dromius | 44.1 | Middle Eocene | Baltic | [86] |
ν | Galerita | 49.0 - 52.0 | Early Eocene | Green River CO | [89] |
ξ | Platynus | 44.1 | Middle Eocene | Baltic | [86] |
ο | Chlaenius | 44.1 | Middle Eocene | Baltic | [86] |
π | Plochionus | 34.9 - 38.0 | Late Eocene | Florissant CO | [89] |
ρ | Pterostichus | 44.1 | Middle Eocene | Baltic | |
σ | Badister | 5.3 - 7.1 | Late Miocene | Oeningen | [91] |
τ | Stenolophus | 34.9 - 38.0 | Late Eocene | Florissant CO | |
ν | Syntomus | 44.1 | Middle Eocene | Baltic | [86] |
ϕ | Helluonini | 44.1 | Middle Eocene | Baltic | [86] |
χ | Harpalini | 61.0 - 65.0 | Early Paleocene | Staratschin cap | [92] |
ψ | Pterostichini | 80 - 90 | Late Cretaceous | Orapa | |
ω | Panagaeus | 23.8 - 28.5 | Late Oligocene | Aix-en-Provance | [95] |
φ | Odacanthini | 34.9 - 38.0 | Late Eocene | Florissant CO | |
ς | Lebia | 44.1 | Middle Eocene | Baltic | [86] |