Table 3 Taxonomy of subfamily Leuciscinae based on the phylogenetic results obtained in this study and lineages that need further review.

N. crysoleucas [152]

N. crysoleucas

Genus Alburnus (Lineage II)

Al. alburnus [2]; Al. arborella [2]; Al. belvica [2]; Al. demiri [153]; Al. escherichii [154]; Al. filippii [154]; Al. hohenackeri [2]; Al. macedonicus [2]; Al. mento [2]; Al. orontis [154]; Al. sarmaticus [2]; Al. thessalicus [2]

Without changes

Alburnus populations from Kizilimark R.

Alburnus sp. 1

Alburnus populations from Stryrmon R.

Alburnus sp. 2

Al. sellal [154]; Al. baliki [155]; Al. kotschyi [156]; Al. orontis [154]

A new genus should be described for these species to avoid the paraphyletic status of Alburnus (Type species: A. alburnus)

Genus Anaecypris (Lineage II)

An. hispanica [2]

An. Hispanica

Genus Leucaspius (Lineage II)

L. delineatus [2]

L. delineatus

Genus Pseudophoxinus (Lineage II)

Ps. punicus [157]

A new genus should be described for this species because real Pseudophoxinus (type species P. zeregi) belong to a different lineage

Genus Scardinius (Lineage III)

Sc. acarnanicus [2]; Sc. erythrophthalmus [2]; Sc. graecus [2]; Sc. hesperidicus [2]; Sc. plotizza [2]; Sc. scardafa [2]

Without changes

Genus Tropidophoxinellus (Lineage IV)

Genus Tropidophoxinellus

Tr. hellenicus [2]; Tr. spartiaticus [2]

Without Changes

Genus Pseudophoxinellus

Ps. callensis [77]

Tr. callensis

Ps. chaignoni [77]

Tr. chaignoni

Genus Petroleuciscus (Lineage V)

Without changes

Pe. borysthenicus [2], Pe. smyrnaeus [2]

Genus Petroleuciscus

Genus Squalius

Sq. aphipsi [2]

Pe. aphipsi

Necessary to analyze all species considered as belonging to Petroleuciscus (P. kurui, P. squaliusculus, P. ulanus)

Genus Ladigesocypris (Lineage V)

Genus Squalius

La. ghigii [2]

La. irideus [2]

Sq. ghigii

Sq. irideus

Genus Squalius (Lineage V)

Sq. albus [2]; Sq. aradensis [2]; Sq. carolitertii [2]; Sq. castellanus [158]; Sq. cephalus [2]; Sq. illyricus [2]; Sq. keadicus [2]; Sq. kosswigi [159]; Sq. kottelati [159]; Sq. laietanus [2]; Sq. lepidus [2]; Sq. lucumonis [2]; Sq. malacitanus [2]; Sq. microlepis [2]; Sq. moreoticus [2]; Sq. orientalis [2]; Sq. orpheus [2]; Sq. pamvoticus [2]; Sq. peloponensis [2]; Sq. prespensis [2]; Sq. pyrenaicus [2]; Sq. squalus [2]; Sq. svallize [2]; Sq. tenellus [2]; Sq. torgalensis [2]; Sq. valentinus [2]; Sq. vardarense [2]; Sq. zrmanjae [2]

Without changes

Squalius populations from Southern Spain

Squalius sp

Squalius populations from Aksheir L. Turkey

Squalius sp_Euphrates

Squalius populations from Euphrates drainage

Squalius sp_Aksheir

Squalius populations from Rama L. BiH

Squalius aff. vardarensis

Squalius populations from Manikiotico R.

Squalius cf. orpheus

Genus Achondrostoma (Lineage VI)

Achondrostoma

Ach. arcasii [2]; Ach. occidentale [2]; Ach. salmantinum [158]; Ach. oligolepis [2]

Achondrostoma populations from NE Spain

Without changes

Achondrostoma sp

Genus Chondrostoma (Lineage VI)

Ch. angorense [160]; Ch. cyri [160]; Ch. holmwoodii [161]; Ch. knerii [2]; Ch. meandrense [160]; Ch. nasus [2]; Ch. oxyrhynchum [2]; Ch. phoxinus [2]; Ch. prespensis [2]; Ch. regium [160]; Ch. soetta [2]; Ch. vardarense [2]

Ch. olisiponensis [162]

Without changes

Iberochondrostoma olisiponensis

Genus Iberochondrostoma (Lineage VI)

Ib. almacai [2];; Ib. lemmingii [2]; Ib. lusitanicum [2]; Ib. oretanum [2]

Without changes

Genus Parachondrostoma (Lineage VI)

Pa. arrigonis [2]; Pa. miegii [2]; Pa. toxostoma [2]; Pa. turiense [2]

Without changes

Genus Phoxinellus (Lineage VI)

Ph. alepidotus [2]; Ph. dalmaticus [2]; Ph. pseudalepidotus [2]

Without changes

Genus Protochondrostoma (Lineage VI)

Pr. genei [2]

Without changes

Genus Pseudochondrostoma (Lineage VI)

Pse. duriense [2]; Pse. polylepis [2]; Pse. willkommii [2]

Without changes

Genus Pseudophoxinus (Lineage VI)

Ps. alii [77]; Ps. anatolicus [154]; Ps. antalyae [154]; Ps. battalgilae [154]; Ps. crassus [154]; Ps. elizavetae [77]; Ps. evliyae [163]; Ps. fahrettini [163]; Ps. firati [77]; Ps. kervillei [164]; Ps. ninae [63]; Ps. zekayi [77]; Ps. zeregi [165]

Without changes

Pseudophoxinus populations from Eflatun Pinari spring. Turkey

Pseudophoxinus sp_EflatunPinari

Genus Telestes (Lineage VI)

Te. alfiensis [63]; Te. beoticus [2]; Te. croaticus [2]; Te. fontinalis [2]; Te. metohiensis [2]; Te. montenegrinus [2]; Te. muticellus [2]; Te. pleurobipunctatus [2];

Te. polylepis [2]; Te. souffia [2]; Te. turskyi [2]; Te. ukliva [2]

Telestes populations from Dreznica. Croatia

Without changes

Telestes sp.

Genus Rutilus (Lineage VII)

Ru. aula [2]; Ru. basak [2]; Ru. frisii [2]; Ru. heckeli [2]; Ru. ohridanus [2]; Ru. panosi [2]; Ru. pigus [2]; Ru. prespensis [2]; Ru. rubilio [2]; Ru. rutilus [2]; Ru. ylikiensis [2]

Without changes

Genus Abramis (Lineage VIII)

Ab. brama [2]

Without changes

Genus Acanthobrama (Lineage VIII)

Genus Acanthobrama

Ac. lissneri [166]; Ac. marmid [154]

Without changes

Genus Acanthalburnus (Lineage VIII)

Aca. microlepis [154]

Ac. microlepis

Genus Petroleuciscus (Lineage VIII)

Pe. persidis [167]

Ac. Persidis

Genus Ballerus (Lineage VIII)

Ba. ballerus [2]; Ba. sapa [2]

Without changes

Genus Blicca (Lineage VIII)

Bl. bjoerkna [2]

Without changes

Genus Mirogrex (Lineage VIII)

Mi. terrasanctae [164]

Without changes

Genus Vimba (Lineage VIII)

Genus Vimba

Vi. melanops [2]; Vi. vimba [2]

Without changes

Genus Acanthobrama

Ac. mirabilis [154] (Vi. Vimba by synonymy [6])

V. mirabilis

Genus Alburnoides (Lineage IX)

Alb. bipunctatus [2]; Alb. prespensis [2]; Alb. bipunctatus [2]

Without changes

Genus Leuciscus (Lineage X)

Genus Leuciscus

Le. idus [2]; Le. latus [167]; Le. leuciscus [2]; Le. schdmidti [46]; Le. walecki [168]

Without changes

Genus Aspius (Lineage X)

As. aspius [2];

Le. aspius

As. vorax [2]

Le. vorax

Genus Pseudophoxinus (Lineage XI)

P. egridiri [154]

A new genus should be described for this species because real Pseudophoxinus (type species P. zeregi) belong to a different lineage

Genus Pachychilon (Lineage XII)

Pac. macedonicum [2]; Pac. pictum [2]

Without changes

Genus Delminichthys (Lineage XII)

D. adpersus [2]; D. ghetaldii [2]; D. jadovensis [2]; D. krbavensis [2]

Without changes

Genus Pelasgus (Lineage XIII)

P. laconicus [2]; P. marathonicus [2]; P. minutus [2]; P. prespensis [2]; P. stymphalicus [2]; P. thesproticus [2]

Without changes

Genus Pelecus (Lineage XIV)

P. cultratus [2]

Without changes