Alternative hypotheses for the evolution of bamboo rats. Sister relationships are constrained to be either (a) Olallamys-Dactylomys or (b) Kannabateomys-Dactylomys. These topologies are statistically equivalent to the unconstrained topology in Figure 4 that finds an Olallamys-Kannabateomys relationship and all three genera unresolved in a polytomy (0.41 PP). Each analysis is pruned from results conducted in BEAST on the full 5-gene data set. For symbols, refer to the legend in Figure 4. See Table 4 for additional details on transition timing and polarity.