Timing of tenrec speciation events and Madagascar colonization. Tree topology as in Figure 1. Divergence times were estimated from the concatenated dataset by a Bayesian relaxed molecular clock method, with six time constraints from fossil calibrations (see Material and Methods). One of them, the paenungulate radiation is represented on the chronogram. Black circles indicate the divergence from the non-Malagasy sister group (node 2) and the initial divergence of Malagasy tenrecs (node 3). Standard deviations are indicated by grey bars, and 95% credibility intervals by open bars. The period of a putative land bridge between Madagascar and Africa at 45–26 Mya  is shaded.