Fig. 1

Time-calibrated phylogenies of the (a) laughingthrush/babbler and (b) shortwing/flycatcher clades. The WG Laughingthrushes, Montecincla, did not group with any of the other laughingthrush clades and are sister to a clade of birds traditionally not included in laughingthrushes. The WG Shortwings, Sholicola, are not part of shortwings (Brachypteryx or Myiomela) but are sister to a clade of Asian blue flycatchers, the Niltavinae. Generic designations follow Clements Checklist [8]; Handbook of the Birds of the World [22] names are shown in brackets if different. Node bars show 95% HPD estimates of divergence dates and stars indicate ML bootstrap of 70% or higher and Bayesian posterior probability of 95% or higher. Letters (in red) show ancestral area reconstruction at relevant nodes (I = Peninsular India; H = Himalayas; A = Southeast Asia)